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- Square: Light Attack (Melee Form)/Fire Bullet 1 (Gun Form)
- Triangle: Heavy Attack (Melee Form)/Fire Bullet 2 (Gun Form)
- O: Dash (Melee Form)/Roll (Gun Form)
- X: Jump
- R1: Switch between Melee or Gun Form
- Hold L1: Lock On/Aiming Reticle (Some gun types can lock on after bringing up the reticle)
- Tilt R3: Swap Lock On Targets
- Hold Triangle: Charge Devour (Melee Only)
- R1 + Triangle: Quick/Air Devour (Melee Only)
- R1 + O: Guard (Melee Only)
- R1 + Square: Special Melee Action (Melee Only)
- R1 + X: Melee to Gun Action (Melee Only)
- *The type of action can be changed in the Options menu
- R1 + O: Special Gun Action (Gun Only)
- R1 + Triangle: Fire Link Burst Shot (Gun Only)
- R2 + X: Dive
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